Знакомство с эволюционной генетикой. Гаевский Н.А. - 24 стр.

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ВУЗ: 

СФУ | Красноярск

Составители: 

• Гаевский Н.А.

Рубрика: 

• Основы генетики

47

>jm]bfih^oh^hffh`_l[ulvihkljh_gb_g_keh`guoh^ghehdmkguofZl_fZlb

-

q_kdbofh^_e_coZjZdl_jbamxsbobaf_g_gb_qZklhludhgljhebjm_fh]hZee_eyaZh^gh

ihdhe_gb_Ijb wlhflj_[m_lky^hims_gb_qlh ^_ckl\b_w\hexpbhgguonZdlhjh\Z^

-

^blb\ghGZijbf_jijbkhq_lZgbbhl[hjZbfmlZpbhggh]hijhp_kkZh[s__baf_g_gb_

qZklhluZee_eyjZ\ghkmff_baf_g_gbcqZklhl\dZ`^hfhl^_evghfijhp_kk_

∆

q

= ∆q + ∆q

(73)

∆

q

.

= pq[q(W

2

-W

1

) + p(W

1

- W

o

)]/W

s

+ pu - qv, (74)

]^_XbYkdhjhklbijyfh]hbh[jZlgh]hfmlbjh\Zgby

Ihkljh_gb_ ]jZnbq_kdbo aZ\bkbfhkl_c ∆q

 hlTiha\hey_l \ jy^_ kemqZ_\ \u-

y\blv^hihegbl_evgu_lhqdbjZ\gh\_kbybebgZclbbaf_g_gbyiheh`_gby^eym`_ km-

s_kl\mxsbo

=_g_lbq_kdZy^bnn_j_gpbZpby\ijhp_kk_\b^hh[jZah\Zgby

H^gbfbaijhy\e_gbcijhp_kkZw\hexpbby\ey_lky\gmljbbf_`\b^h\Zybaf_g-

qb\hklvJZaebqby\n_ghlbibq_kdboijbagZdZohkgh\ZgugZkms_kl\mxsbo]_ghlbib-

q_kdbojZaebqbyoIhke_^gb_h[gZjm`b\Zxlk_[ydh]^Zm^Z_lkyihemqblv\lhjh_ihdh-

e_gb_ihlhfdh\ ijbkdj_sb\Zgbb[ebadhjh^kl\_gguo\b^h\H^gZdh \ jZfdZo f_g^_-

e_\kdhc]_g_lbdbg_\hafh`ghhl\_lblvgZ\hijhkdZdh\Zkl_i_gv]_g_lbq_kdbojZaeb-

qbcf_`^mihimeypbyfbh^gh]h\b^Zbebf_`^mjZaebqgufb\b^ZfbK^jm]hcklhjh-

gu hl\_lb\ gZ wlhl \hijhk \ jZfdZo fhghnbe_lbq_kdhc dhgp_ipbb hj]Zgbaf_gghc

w\hexpbbfh`ghg_lhevdhmklZgh\blvkl_i_gvjh^kl\ZghbkjZ\gblvw\hexpbhggu_

jZkklhygbyhl^_eyxsb_ij_^klZ\bl_e_c^\mojZaebqguo\b^h\^jm]hl^jm]Zbhlbo

h[s_]hij_^dZ

<gZklhys__\j_fykms_kl\m_lg_kdhevdhf_lh^h\^eyhp_gdb]_g_lbq_kdhc^bn-

n_j_gpbZpbb^\moihimeypbcFh`ghmihfygmlvf_lh^hp_gdblh`^_kl\_gghklb]_gh-

fh\ b [he__ ih^jh[gh hklZgh\blvky gZ f_lh^_ ZgZebaZ we_dljhnhj_lbq_kdbo k\hckl\

[_edh\ IheZ]Zxl qlh dZ`^hfm Zee_ex \ ehdmk_ khhl\_lkl\m_l k\hy bahnhjfZ [_edZ

hlebqZxsZykyhl^jm]hcbahnhjfuk\h_cih^\b`ghklvx\lhdhijh\h^ys_f]_e_

FZkZlhrbG_c[1]ij_^eh`be^bnn_j_gpbjh\Zlv^\_ihimeypbbgZ hkgh\_we_d-

ljhnhj_lbq_kdh]h ZgZebaZ [_edh\ < dZq_kl\_ oZjZdl_jbklbd ]_g_lbq_kdhc ^bnn_j_g-

pbZpbbij_^eh`_gh^\Z iZjZf_ljZ]_g_lbq_kdh_koh^kl\h,dZd ^hey kljmdlmjguo ]_-

gh\dhlhju_b^_glbqgu\h[_boihimeypbyob]_g_lbq_kdh_jZkklhygb_'dZdkj_^g__

qbkehaZf_gZee_e_c\dZ`^hfehdmk_

48

JZkkfhljbfijbf_jhp_gdb]_g_lbq_kdhc^bnn_j_gpbZpbbq_luj_o\b^h\ijbfZ

-

lh\lZ[egZhkgh\_ZgZebaZehdmkh\dh^bjmxsbo [_edbdjh\b<klhe[pZo^ey

dZ`^h]h\b^Zijb\_^_gupbnjh\u_bg^_dkuZee_e_cbl^<kemqZyodh]^Z

ehdmkihebfhjn_g\kdh[dZomdZaZgukhhl\_lkl\mxsb_qZklhluZee_e_c

LZ[ebpZ

Ehdmk RbfiZga_ =hjbeeZ =b[[hg ;Z[mbg

12 3 4 5

1. Ak 96 98 (0.20)

100 (0.80)

92 96

2. Alb 100 100 100 99

3. Aph 100 100 100 100

4. Cer 100 98 98 102

5. Dia 100 85 (0.67)

95 (0.33)

100 (0.67)

108 (0.33)

95 (0.88)

100 (0.12)

6. Est-A 100 101 102 96

7. Est-B 100 100 102 95 (0.17)

96 (0.08)

103 (0.75)

8. G6pd 100 100 102 102

9. Got 100 100 96 96 (0.14)

100 (0.86)

10. Hb 100 100 100 100 (0.92)

102 (0.08)

11. Hpt 105 107 107 107

12. Icd 96 100 100 100 (0.94)

107 (0.06)

13. Lap 100 100 100 100

14. Ldh-

A

96 96 96 96

15. Ldh-

B

100 100 100 100

16. Mdh 100 100 93 (0.62)

100 (0.38)

106

17. 6-Pgd 97 97 (0.15)

105 (0.85)

94 94

18. Pgm-

1

96 (0.12)

100 (0.88)

100 100 94

19. Pgm-

2

100 96 100 102

>eyjZkq_lZ]_g_lbq_kdh]hkoh^kl\Z^\moihimeypbcihh^ghfmehdmkmbkihevamxlke_-

^mxs__\ujZ`_gb_

I

ab

ab

k

ii

ii

=

∑

∑∑

22

, (75)

]^_a

i

bb

i

qZklhluZee_e_ci - ehdmkZ\ihimeypbyo:b<

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